Phacelia, the largest genus in the family Hydrophyllaceae, consists of some 150 to 200 species. The genus is heterogeneous and has undergone several revisions of both infrageneric organization and species classification. The type species is P. secunda J. F. Gmelin, Syst. 2:330. 1791.
In 1789. Jussieu assigned the genus to what is now the family Boraginaceae. Brown (see Edwards, 1817) assigned Phacelia to the order Hydrophylleae. In 1875 Asa Gray united the order Hydrophylleae with the family Hydroleae and formed the family Hydrophyllaceae. He divided the genus Phacelia into seven subgenera based on characteristics of ovules, seeds, and corolla appendages. The subgenera included Euphacelia, Eutoca (formerly a separate genus),
Microrgenetes, Whitlavia, Gymnobythus, Cosmanthus, and Cosmanthoides. In 1890 Phacelia was once more put in the Boraginaceae by Baillon, who did not recognize the family Hydrophyllaceae. In 1913, however, Brand restored the Hydrophyllaceae to family status and wrote the only existing monograph dealing with the entire genus Phacelia. Hedivided the genus into six sections, four of which were very small. The small sections were Whitlavia, Gynmobythus, Microgenetes, and Cosmanthus; the two large ones were Eutoca and Euphacelia. Partial revisions of the genus by several authors followed.
Level of Degree
UNM Biology Department
First Committee Member (Chair)
William Clarence Martin
Second Committee Member
Clarence Clayton Hoff
Third Committee Member
Howard J. Dittmer
Bobisud, Helen M.. "A Study Of The Genus Phacelia In New Mexico." (1967). https://digitalrepository.unm.edu/biol_etds/352